my guess is last journey(Hawaii to USA mainland) wore them out....
these fishes went through system changes quickly...boat journey from collection point(5-10 days),then on collector's hold-up facility(15-30 days),then some 12 hours journey to another system...
most common fishes go through local collection,then wholesellers and then lfs...similar changes..but these fishes seem to be more sensitive..
even the smallest specimen of this expedition died in US shore in few days...seems like smaller the better is not applicable for it..being small actually acted as a disadvantage...without eating died quickly...